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Etting a target FDR threshold of 1 at the peptide level. Mass spectrometric analysis resulted in identification of a total of 20,783 peptides. After removing peptides not labelled with all the four labels (n = 212) and those (n = 1968) shared between multiple proteins, 18,603 peptides were considered for identification of proteins. The labelling efficiency was thus 99 . Relative quantitation of p

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